Content
Topic Introduction
The insects and their kin (the Hexapoda) constitute a subphylum within the phylum Arthropoda. They are the most diverse group of animals on Earth. They are the only invertebrates that can fly, and the only ones to undergo indirect development or complete metamorphosis. They comprise the class Insecta, as well as three small, closely-related wingless groups: the Collembola, Protura and Diplura. All hexapods have a three-part bodyplan, with a head, a 3-segmented thorax and an 11-segmented abdomen. They have three pairs of thoracic legs, with one pair of legs per segment.
Hexapods evolved on land, and have occupied almost every known habitat. Some have moved into aquatic habitats, including the marine environment. There are an unknown number of species, with some estimates being over 100 million species. To date, approximately a million species have been described, with many new species (5000–7000) being described every year. That compares to a total of about 6500 mammals species described to date.
There are so many different types of insects that it is impossible to undertake a thorough review in a course such as this. Instead, we provide an introduction that summarizes the different groups and provides an illustration of each, and then we examine three particular groups: the Odonata (dragonflies and damselflies), Coleoptera (beetles), Hymenoptera (ants, bees and wasps) and Lepidoptera (moths and butterflies). With this last group (Lepidoptera), we focus on the butterflies, which are much better known than the moths. This was a hard choice. There are many other important groups (such as the Orthoptera (locusts, katydids, crickets and grasshoppers), Hemiptera (true bugs), Thysanoptera (thrips), and Diptera (true flies, mosquitoes and gnats), as well as some fascinating groups such as the Mantodea (mantises), Phasmida (stick and leaf insects), and Neuroptera (lacewings, ant lions, mantisflies and owlflies). In our examination of the Hymenoptera, we have included the termites, even though they belong to a different order of insects (Blattodea).
By the end of this topic, you should be able to:
- Describe the main orders of insects.
- Assess different explanations for the diversity of the insects.
- Explain why the insects are important for the ecology of tropical forests.
- Distinguish between dragonflies and damselflies.
- Explain why some regions have richer odonate faunas than others.
- Describe some of the most important beetle families found in tropical forests.
- Demonstrate their ecological roles in forests.
- Give examples of different types of ants and their life histories.
- Contrast the roles that different types of bees play in pollination.
- Evaluate the ecological significance of ant gardens and ant plants.
- Explain the difference between butterflies and moths.
- Give examples of each of the five butterfly families.
- Contrast the life histories of butterflies in tropical moist and seasonally dry forests.
Topic Readings
Required Readings
- Ghazoul, J. and Sheil, D. 2010. Tropical Rain Forest Ecology, Diversity, and Conservation. Oxford, Oxford University Press, 516 pp. Chapter 5.1.
Optional Readings
General
- Brusca, R.C., Moore, W. and Shuster, S.M. 2016. Invertebrates. 3rd Sunderland, Massachusetts, USA: Sinauer Associates. Chapters 22.
- Hanson, P.E. 2011. Insect diversity in seasonally dry tropical forests. In: Dirzo, R., Young, H.S., Mooney, H.A., and Ceballos, G. (eds.) Seasonally Dry Tropical Forests: Ecology and Conservation. Washington DC, USA: Island Press. pp. 71-84.
- Roubik, D.W. 1992. Ecology and Natural History of Tropical Bees. Cambridge, UK: Cambridge University Press. 514 pp.
Further Readings
Field Guides
Australia and New Guinea
- Braby, M.F. 2004. The Complete Field Guide to Butterflies of Australia. Collingwood, Victoria, Australia: CSIRO Publishing. 339 pp.
- Brock, P.D. and Hasenpusch, J.W. 2009. The Complete Field Guide to Stick and Leaf Insects of Australia. Collingwood, Victoria, Australia: CSIRO Publishing. 204 pp.
- Golding, M.R. 2016. A Guide to Australian Tiger Beetles. Mark R Golding. 188 pp.
- Hangay, G. and de Keyzer, R. 2017. A Guide to Stag Beetles of Australia. Collingwood, Victoria, Australia: CSIRO. 245 pp.
- Hangay, G. and Zborowski, P. 2010. A Guide to the Beetles of Australia. Collingwood, Victoria, Australia: CSIRO Publishing. 238 pp.
- Houston, T. 2018. A Guide to Native Bees of Australia. Collingwood, Victoria, Australia: CSIRO Publishing. 272 pp.
- Lawrence, J.F. and Ślipiński, S.A. 2013. Australian Beetles, Volume 1. Morphology, Classification and Keys. Collingwood, Victoria, Australia: CSIRO. 561 pp.
- Michalski, J. (ed.) A Manual for the Identification of the Dragonflies and Damselflies of New Guinea, Maluku, and the Solomon Islands. John Michalski. 579 pp.
- Patrick, B., Patrick, H. 2012. Butterflies of the South Pacific. Otago University Press. 240 pp.
- Rentz, D. 2010. A Guide to the Katydids of Australia. Collingwood, Victoria, Australia: CSIRO Publishing. 214 pp.
- Rentz, D. 2014. A Guide to the Cockroaches of Australia. Collingwood, Victoria, Australia: CSIRO. 318 pp.
- Rentz, D. and Su, Y.N. 2019. A Guide to Crickets of Australia. Collingwood, Victoria, Australia: CSIRO Publishing. 398 pp.
- Ślipiński, S.A. and Escalona, H.E. Australian Longhorn Beetles (Coleoptera: Cerambycidae), Volume 1. Introduction and Subfamily Lamiinae. Collingwood, Victoria, Australia: CSIRO. 484 pp.
- Ślipiński, S.A. and Escalona, H.E. 2016. Australian Longhorn Beetles (Coleoptera: Cerambycidae), Volume 2. Subfamily Cerambycinae. Collingwood, Victoria, Australia: CSIRO. 613 pp.
- Ślipiński, S.A. and Lawrence, J.F. (eds.) 2019. Australian Beetles, Volume 2. Archostemata, Myxophaga, Adephaga, Polyphaga (Part). Collingwood, Victoria, Australia: CSIRO. 784 pp.
- Theischinger, G. and Hawking, J. 2006. The Complete Field Guide to Dragonflies of Australia. Collingwood, Victoria, Australia: CSIRO Publishing. 366 pp.
- Webb, C., Doggett, S.L. and Russell, R.C. 2016. A Guide to Mosquitoes of Australia. Collingwood, Victoria, Australia: CSIRO. 204 pp.
- Zborowski, P. and Edwards, T. 2007. A Guide to Australian Moths. Collingwood, Victroia, Australia: CSIRO Publishing. 214 pp.
- Zborowski, P. and Storey, R. 2010. A Field Guide to Insects of Australia. 3rd Sidney, Australia: Reed New Holland. 328 pp.
Southeast and South Asia
- Assam State Biodiversity Board. 2015. Common Spiders from Selected Protected Areas of Upper Assam. Guwahati, Assam, India: Assam State Biodiversity Board. 192 pp.
- Bhakare, M. and Ogale, H. 2018. A Guide to Butterflies of Western Ghats (India). Milind Bhakare. 496 pp.
- Bosuang, S., Audibert, C., Porion, T., and Chan, C.L. 2017. A Guide to Lanternflies of Borneo. Kota Kinabalu, Malaysia: Natural History Publications Borneo. 120 pp.
- Bosuang, S., Chung, A.Y.C., and Chan, C.L. 2017. A Guide to Beetles of Borneo. Kota Kinabalu, Malaysia: Natural History Publications Borneo. 244 pp.
- Brock, P.D. 1999. Stick and Leaf Insects of Peninsular Malaysia and Singapore. Malaysian Nature Society. 222 pp.
- Hardy, P.B. and Lawrence, J.M. 2017. Field Guide to Butterflies of The Philippines. Siri Scientific Press. 448 pp.
- Jayasinghe, H.D. 2015. Common Butterflies of Sri Lanka. Merrill J Frenando and Sons. 176 pp.
- Kehimkar, I. 2016. Butterflies of India. Mumbai, India: Bombay Natural History Society. 505 pp.
- Khoon, K.S. 2015. A Field Guide to the Butterflies of Singapore. Marshall Cavendish. 376 pp.
- Mitra, T.R. 2006. Handbook of Common Indian Dragonflies (Insecta: Odonata). Zoological Survey of India. 136 pp.
- Mita, A. and Mitra, B. 2009. Pictorial Handbook on Dragon and Damselflies (Odonata: Insecta) on Mangroves of Sunderbans. Zoological Survey of India. 56 pp.
- Nair, M.V. 2011. Dragonflies and Damselflies of Orissa and Eastern India. Orissa Wildlife Organization. 252 pp.
- Neog, S. 2015. Butterflies of Gibbon Wildlife Sanctuary. Bhabani Books. 112 pp.
- Orr, A.G. 2003. A Guide to the Dragonflies of Borneo. Their Identification and Biology. Kota Kinabalu, Malaysia: Natural History Publications Borneo. 195 pp.
- Phillipps, H. 2020. A Naturalist’s Guide to the Butterflies of Borneo. John Beaufoy Publishing. 176 pp.
- Rangnekar, P. 2007. A Photographic Guide to Butterflies of Goa. Broadway Publishing House. 66 pp.
- Reels, G. and Zhang, H. 2015. A Field Guide to the Dragonflies of Hainan. Kadoorie Farm and Botanic Garden. 463 pp.
- Saini, M., Raina, R.H. and Ghator, H.S. 2015. Indian Bumblebees. Bishen Singh Mahendra Pal Singh. 248 pp.
- Seow-Chen, F. 2005. A Pocket Guide: Phasmids of Peninsular Malaysia and Singapore. Kota Kinabalu, Malaysia: Natural History Publications Borneo.
- Seow-Choen, F. 2018. A Taxonomic Guide to the Stick Insects of Sumatra, Volume 1. Kota Kinabalu, Malaysia: Natural History Publications Borneo. 703 pp.
- Sharma, G. 2015. Pictorial Handbook on Damselflies and Dragonflies (Odonata: Insecta) of Rajasthan. Zoological Survey of India. 266 pp.
- Shubhalaxmi, V. 2018. Birdwing Field Guide to Indian Moths. Birdwing Publishers. 460 pp.
- De Silva Wijeyeratne, G. 2018. A Naturalist’s Guide to the Butterflies and Dragonflies of Sri Lanka. John Beaufoy Publishing. 176 pp.
- Smetacek, P. 2016. A Naturalist’s Guide to the Butterflies of India. John Beaufoy Publishing. 176 pp.
- Sondhi, S. and Kunte, K. 2018. Butterflies of Uttarakhand, A Field Guide. Bishen Singh Mahendra Pal Singh. 310 pp.
- Sumanapala, A.P. 2017. A Field Guide to the Dragonflies and Damselflies of Sri Lanka. Dilmah Conservation. 174 pp.
- Sutton, S., Barlow, H., and Whitaker, T. 2015. A Preliminary Guide to the Pyralid Moths of Borneo, Part 1: Thyridoidea and Pyraloidea. Koata Kinabalu, Malaysia: Natural History Publications Borneo. 89 pp.
- Tze-Wai, T., Ka-King, L., Kwan, B.S.P., Wu, K.K.Y., Tang, S.S.H., Do, I.W.Y., Cheng, J.C.Y., Yuen, E.F.M., Yu-Man, T. Wing-Leun, H., Agriculture, Fisheries and Conservation Department 2011. The Dragonflies of Hong Kong. Huayu Book Trade Co. 368 pp.
- Van der Poorten, G.M. and Van der Poorten, N. 2018. Field Guide to the Butterflies of Sri Lanka. Lepodon Books. 250 pp.
- Vedam, A.P. 2011. Butterflies of India. Vedam Books International. 184 pp.
Africa
- Braack, L.E.O. 1991. Field Guide to Insects of the Kruger National Park. Cape Town, South Africa: Struik. 158 pp.
- Fisher, B.L. and Bolton, B. 2016. Ants of Africa and Madagascar. A Guide to the Genera. University of California Press. 539 pp.
- Lawrence, J.M. 2014. Field Guide to Butterflies of Seychelles. Their Natural History and Conservation. Siri Scientific Press. 125 pp.
- Martiré, D. 2010. Les Libellules et Éphémères de La Réunion. Biotope. 72 pp.
- Penney, D. 2009. Field Guide to Butterflies of the Gambia, West Africa. Siri Scientific Press. 80 pp.
- Picker, M., Griffiths, C. and Weaving, A. 2004. Field Guide to Insects of South Africa. Cape Town, South Africa: Struik. 444 pp.
- Slingsby, P. 2017. Ants of Southern Africa. The Ant Book for All. Slingsby Maps. 247 pp.
- Tarboton, W., Tarboton, M. 2019. A Guide to the Dragonflies & Damselflies of South Africa. 2nd Penguin Random House South Africa.
- Woodhall, S. 2005. Field Guide to the Butterflies of South Africa. London, UK: Bloomsbury Publishing. 352 pp.
Madagascar
- Fisher, B.L. and Bolton, B. 2016. Ants of Africa and Madagascar. A Guide to the Genera. University of California Press. 539 pp.
- Fisher, B.L. and Peeters, C. 2019. Ants of Madagascar: A Guide to the 62 Genera. Association Vahatra. 260 pp.
Central and South America
- Albert, R. 2010. Butterflies of the Golfo Dulce Region, Costa Rica. Tropical Field Station La Gamba. 76 pp.
- Cândido de Almeida, A. and Freitas, A.V.L. 2012. Lepidoptera: Borboletas e Mariposas do Brasil. Grupo Direcional. 208 pp.
- Catling, P.M. and Kostiuk, B. 2015. Butterflies of St. Kitts and Nevis and the Northeastern Leeward Islands, West Indies. Nevis Historical and Conservation Society. 98 pp.
- Debrot, A.O. and Miller, J.Y. 2017. Butterflies and Moths of Curaçao, Aruba and Bonaire. CARIB Publishing. 102 pp.
- Dett, A. 2017. Moths of Costa Rica’s Rainforest. Benteli Verlag AG. 336 pp.
- Fontana, P., Buzzetti, F.M. and Mariño-Pérez, R. 2008. Grasshoppers, Locusts, Crickets and Katydids of Mexico. WBA Handbooks, Voluime 1. World Biodiversity Association. 272 pp.
- Fontana, P., Buzzetti, F.M., Mariño-Pérez, R., Castellanos-Vargas, I., Monge-Rodriguez, S., and Cano-Santana Z. 2017. Orthopterans of Oaxaca: Photographic Field Guide. World Biodiversity Association. 209 pp.
- Garwood, K. and Lehman, R. 2011. Butterflies of Central America, Volume 1: Swallowtails, Pierids, and Brushfoots. Kim Garwood. 304 pp.
- Garwood, K. and Lehman, R. 2012. Butterflies of Central America, Volume 2: Lycaenid & Riodinidae: The Hairstreaks and Metalmarks. Kim Garwood. 235 pp.
- Garwood, K. and Lehman, R. 2012. Butterflies of Central America, Volume 3: Hesperiidae, The Skippers. A Photographic Checklist of Common Species. Kim Garwood. 287 pp.
- Glassberg, J. 2018. A Swift Guide to the Butterflies of Mexico and Central America. Princeton, New Jersey, USA: Princeton University Press. 304 pp.
- Gowin, E. 2017. Mariposas Nocturnas: Moths of Central and South America, A Study in Beauty and Diversity. Princeton, New Jersey, USA: Princeton University Press. 144 pp.
- Hanson, P.E. 2016. Insects and Other Arthropods of Tropical America. Comstock Publishing Associates. 375 pp.
- Henderson, C.L. 2010. Butterflies, Moths, and Other Invertebrates of Costa Rica. A Field Guide. University of Texas Press. 173 pp.
- Lingafelter, S.W., Wappes, J.E., and Arias, J.L. 2017. Photographic Guide to Longhorned Beetles of Bolivia. Washington, DC, USA: Smithsonian Institution Scholarly Press. 260 pp.
- Kompier, T. 2015. A Guide to the Dragonflies and Damselflies of the Serra dos Orgaos, South-eastern Brazil / Guia dos Anisoptera e Zygoptera da Serra dos Órgãos, Sudeste do Brasil. REGUA. 379 pp.
- Martin, M. 2011. A Guide to the Hawkmoths of the Serra dos Orgaos, Southeastern Brazil / Guia dos Sphingidae da Serra dos Órgãos, Sudeste do Brasil. 143 pp.
- Vieira, R.S., Motta, C., Agra, D.B., Lopes, L.M. and Cezar, K.F.S. 2014. Guia Ilustrado de Borboletas da Reserva Florestal Adolpho Ducke. Instituto de Pesquisas da Amazonia. 132 pp.
Self-check Quiz
After viewing topic presentations, take this self-check quiz to check your own progress. This quiz is designed to see how well you remember some of the more important issues presented in the topic. There are five questions, and each has a potential of 4 answers, only one of which is correct. When you are ready, click the link Self-check Quiz 5.4 to start taking the quiz.
Pre-readings and Discussion (For certification)
Once you have read Section 5.1 of Ghazoul, J. and Sheil, D. 2010. Tropical Rain Forest Ecology, Diversity, and Conservation, post one thought provoking question related to the material that you have read, and answer/respond to at least one other student’s question on the discussion board Online Discussion Module 5.
Graded Assignment (For certification)
Six Sentence Answer #10
If you are not familiar with the Six Sentence Answer (6SA) format, please refer to the course pages on graded assessments to see details of the 6SA exercise. You should not attempt to do this assessment until you have read and fully understood the requirements.
The Case
So far, we presented the situation that you are a member of a major international forest consultancy firm that specializes in doing surveys of forests around the world. You have access to field teams and the latest surveying equipment, including LiDAR. Your company has taken on a contract with the UN-REDD (United Nations Collaborative Programme on Reducing Emissions from Deforestation and Forest Degradation in Developing Countries) to survey the forests on a very large island in South-East Asia, and you not only have the full cooperation of the government but they are also asking your company for advice on a range of other issues.
You have already produced a map of the island’s forests, and assessed the growing stock. Your field teams have done on-the-ground surveys to develop the allometric equations that you needed to produce estimates of the carbon stored in different types of forest on the island. You have found major differences, as you might expect, between the moist tropical forests on the west of the island and the dry forests in the rain shadow of the volcano on the east of the island. You have identified that the trees in the tropical moist forest are much smaller than those on the island of Borneo, located only a few hundred kilometres to the west and you have found a plausible explanation for this.
The government has indicated that it is interested in developing ecotourism on the island and are building an interpretative walk through a section of rain forest on the west of the island. They are still at the initial stages of planning, and have asked your company to help them. You have already indicated how you would prioritize the botanic features along the walk and how the plants got there, and how you would also develop an interpretative sign for the mammals presents on the island.
In order to save money, the development company working on the tourism infrastructure, such as the large hotels, has brought in a significant number of migrant workers from somewhere in Southeast Asia. A number of workers have fallen sick, and one has died. Doctors in the only hospital on the island have indicated that they have, for the first time on the island, recorded a protozoan parasite called Plasmodium falciparum in blood samples taken from the sick workers. Your company has been tasked with finding a solution to this problem. Using the 6SA format, please indicate what you would recommend.
Remember to follow the structure of the 6SA outlined in the section on graded assessments.
Topic Summary
In this final topic of the course, we have had a very brief look at the most diverse of all the animal groups on Earth: the hexapods. These mostly comprise the insects, which presently occupy almost every habitat available to them. They have been incredibly successful, not only in terms of their diversity, but also in terms of the numbers of individuals within each species. Insects are believed to make up 25–45% of the animal biomass of tropical forests, an extraordinary feat given the presence of some very large vertebrates, such as elephants and rhinoceroses.
Unfortunately, most insect species remain undescribed. We really have no idea how many species currently exist: some models such that it may be as many as 100 million. This presents us with a huge challenge when talking about the biodiversity of tropical forests. When we do, we often are referring only to the vertebrate or plant diversity, as statements about the total insect diversity are close to meaningless. We know some groups better than others: these tend to be the groups that are more easily observed, such as the spiders, butterflies, dragon- and damselflies, and beetles. However, new species continue to be described in large numbers, and this will continue into the foreseeable future.
Topic Self-review (For self learning)
To review what you have learned in the topic, you are encouraged to use the following reflection questions as a study guide to do a self-review for the topic:
- What are the main orders of insects?
- What do you think is the most likely explanation for the diversity of the insects?
- Why are insects so important for the ecology of tropical forests?
- What are the differences between dragonflies and damselflies?
- Why do some regions have richer odonate faunas than others?
- What are the most important beetle families found in tropical forests?
- What ecological roles do beetles play in tropical forests?
- Can you give examples of different types of ants and their life histories?
- What roles do the different types of bees play in pollination?
- What, if any, is the ecological significance of ant gardens and ant plants?
- Explain the difference between butterflies and moths?
- Can you give examples of each of the five butterfly families?
- How do the life histories of butterflies in tropical moist and seasonally dry forests compare?
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